Leaf blade usually lanceolate to ovate, rarely linear or subulate, herbaceous, rarely leathery.

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Leaves simple, opposite, rarely whorled or alternate, pinnately veined; stipules absent or rarely present.

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Flowers small, bisexual or rarely unisexual, actinomorphic, rarely zygomorphic.

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Flowers solitary, sessile, rarely clustered and stalked, bisexual, rarely unisexual, actinomorphic or occasionally zygomorphic.

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Seeds uniseriate or rarely biseriate, winged, margined, or wingless, oblong, plump or flattened; seed coat minutely reticulate, mucilaginous when wetted; cotyledons incumbent or rarely accumbent.

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Seeds biseriate or rarely uniseriate, wingless or narrowly margined, oblong, plump or slightly flattened; seed coat minutely reticulate, copiously mucilaginous when wetted; cotyledons incumbent or rarely accumbent.

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Petals 5 or rarely more, white or rarely pale yellow, basally slightly connate and adnate to stamens.

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Shrubs (sometimes scrambling or rarely somewhat climbing) or rarely small trees, monoclinous or andromonoecious, armed or unarmed, apparently without oil glands.

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Fruit juicy or dry, naked, scaly, hairy, bristly, or spiny, indehiscent or dehiscent, when juicy then pulp derived from often deliquescent funicles. Seeds usually numerous, often arillate or strophiolate; embryo curved or rarely straight; endosperm present or absent; cotyledons reduced or vestigial, rarely leaflike.

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Leaves simple, usually alternate, rarely opposite or verticillate, sometimes crowded at apices of branches; stipules usually small and caducous, sometimes larger, leaflike and persistent, rarely absent; petiole generally present, sometimes with apex 2-glandular and/or with additional glands along petiole length; leaf blade usually pinnate-veined, sometimes 3-5-veined from base or palmate-veined, with or without pellucid dots or lines, sometimes with a pair of glands at junction of blade and petiole, margin entire or toothed, teeth glandular or not. Inflorescences axillary, terminal, or cauliflorous, of various forms: racemose, spicate, cymose, corymbose, or paniculate, sometimes flowers fasciculate, or solitary; pedicels often articulate; bracts and bracteoles usually small to minute.

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