Nectar scales oblong-cuneate to linear-oblong, apex rounded or subemarginate.

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Fruiting pedicels clavate at apex; fruit oblong, elliptic, oblong-oblanceolate, or elliptic-obovate, glabrous or pubescent with slender trichomes, apex acute, obtuse, rounded, or rarely subemarginate.

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Calyx 2-3 mm; lobes suborbicular, rounded, margin submembranous.

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Stipules gradually deciduous, triangular, 1-2 mm, membranous; petiole often unequal in length, 0.5-7 cm; leaf blade dark green or green blue, ovate, lanceolate, or obovate-oblong, asymmetric, unequal or equal in size, 1-15 × 0.6-5 cm, thinly papery or submembranous, 3(or 5)-veined, lateral veins many, irregularly reticulate, external secondary veins anastomosing by margin, sometimes inconspicuous, cystoliths finely fusiform, conspicuous adaxially, base obliquely rounded or subcordate, sometimes cuneate, margin entire or sometimes undulate, apex shortly or long caudate-acuminate. Inflorescences solitary; male inflorescence a cymose panicle or racemelike, often longer than leaves; female cymes as long as or shorter than leaves, pedunculate.

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Flowers green; calyx lobes rounded or subreniform, ca. 0.5 × 0.75-1 mm; petals elliptic or broadly ovate, 1.25-1.5 × ca. 1 mm, reflexed at anthesis, obtuse or slightly acute.

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Seed scales triangular-ovate, ovate, or subrhombic, usually densely purplish brown hairy abaxially, apex rounded

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Basal petioles 1.5–5 cm, sheaths long, broad; blade oblong, 2–4 × 1–2 cm, pinnate; pinnae sessile,(3–) 4–6 pairs, remote; ultimate segments long-ovate to suborbicular, 5–10 × 3–8 mm, base subtruncate or rounded, entire or apex shallowly 1–3-toothed, sometimes 3–4-crenate; terminal pinnae obovate or orbicular, base cuneate, apex more deeply 3-lobed.

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Features of male owstoni are: dark green back; bi-coloured ear coverts; white spur on tertials; yellow supercilium reaching the nostril; blue-grey on some median and lesser coverts; rather rounded, short primaries; lack of orange on throat and rump.

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Stems terete when mature; leaves ovate to elliptic or obovate, subacute to rounded

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Mm. Stamens as long as petals; anthers rounded with 2 thecae.

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