Shrubs or trees. Leaves alternate, petiolate; stipules deciduous or evergreen, small, pinnately veined; leaf blade simple, elliptic to lanceolate, secondary veins spreading or curved, margin glandular dentate or spinose dentate, rarely crenate or entire.

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Plants annual. Petiole 5-10 cm; leaf blade palmately (3 or)5-9-lobed, sometimes simple, 7-10 × 7-10 cm, papery, abaxially with rigid spinulose hairs on veins, adaxially pubescent but not densely so, base cordate; lobes ovate-triangular, margin serrate.

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Leaf blade thickly papery, linear-lanceolate, often strongly rugose adaxially, apex tapering to acuminate; female spikes of congested glomerules, simple, nodding; achene long stipitate.

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Such plants usually have one or more domatia, cavities in stems, leaves or roods that house ants. These may be elaborate systems of cavities with many chambers, e.g. Myrmecodia, or simple hollows in internodes, leaf veins, petioles or stipular spines, e.g. whistling thorns.

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Thalli A simple plant body showing no differentiation into root, leaf, and stem and lacking a true vascular system.

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Leaves in spirals, odd-pinnate, trifoliolate, or simple, when compound lateral leaflets opposite; leaf blade or leaflet blades with margin entire or sparsely crenate.

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Stems unarmed, sometimes with conspicuous lenticels, or bark sometimes shredding; branches often swollen at 3-7-lacunar nodes; pith continuous or interrupted by diaphragms at nodes; tendrils simple, bifurcate to trifurcate, or 4-12-branched, usually leaf-opposed, rarely tendrils absent.

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Indumentum of simple hairs, glandular hairs, or multicellular hairs secreting calcium oxalate and forming scales or present beneath cuticle and making leaf blade surface verruculose and sometimes translucent dotted.

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Shrubs, usually aerial hemiparasites on other seed plants, often spreading along host by runners, more rarely terrestrial root-parasitic shrubs or trees, nodes not articulated, glabrous or hairy, hairs often stellate or verticillate. Leaves opposite or alternate, stipules absent; petiole often indistinct; leaf blade simple, usually pinnately veined, margin entire.

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Sometimes armed with stinging hairs; epidermal cells of leaves, sometimes stems, perianths mostly with prominent cystoliths punctiform to linear; Leaves alternate or opposite, stipules present, rarely absent; leaf blade simple. Inflorescences cymose, paniculate, racemose, spicate, or cluster-capitate, usually formed from glomerules, sometimes crowded on common enlarged cuplike or discoid receptacle, rarely reduced into a single flower. Flowers unisexual, rarely bisexual in partial flowers; actinomorphic, very small,(1-)4- or 5-merous, rarely perianth absent in female flowers.

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