Stems 50-150 cm tall, usually branched throughout, densely to very sparsely strigillose, with long suberect red pustulate-based hairs, and glandular hairs on inflorescence.

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Ovary strigillose, not white canescent; seeds with inconspicuous chalazal collar; leaves sublinear to elliptic or lanceolate, 1-4.5 cm

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Capsules 2.5-5 cm, strigillose; pedicels 0.5-1 cm. Seeds light brown, 1.1-1.5 mm, low papillose, with inconspicuous chalazal collar; coma white, detaching easily.

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Sepals 2 or 3; petals 5-12; leaves adaxially not strigillose

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Anthers 4-10 mm; pollen ca. 50% fertile. Ovary densely strigillose; stigma surrounded by anthers.

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Herbs perennial or subshrubs; petiole to 6 or 13 cm; female inflorescences solitary; female perianth tube strigillose on shoulder.

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Inflorescence strigillose and/or villous, but lacking short, erect glandular hairs

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Stems 30-150 cm tall, simple or sparsely branched, sparsely strigillose mixed with glandular and long spreading pustulate-based hairs.

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Leaves alternate; stipules lanceolate, 4-8 mm; petiole 2-11(-13) cm, pubescent or glabrous; leaf blade ovate, elliptic, oblong or subelliptic,(5-)9-21 × 2.5-10 cm, papery, slightly rough, secondary veins 3 or 4 pairs along midvein, abaxial surface strigillose or pubescent on veins, rarely glabrous, adaxial surface sparsely appressed strigillose or glabrous, base broadly cuneate, rounded, or obtuse, margin denticulate, apex acuminate or caudate-acuminate. Glomerules usually unisexual, sometimes bisexual, in the axils of current and fallen leaves along normal leafy stems, 4-10 mm in diam.; male ones on proximal part of flower-bearing branches; female ones on distal part.

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Male flowers 4-merous, sessile; perianth lobes elliptic, ca. 1 mm, strigillose, connate at base. Fruiting perianth rhomboid-obovoid, compressed, ca. 1.2 mm, smooth, strigillose on shoulder, base cuneate or stipitate, apex with short neck, 2-toothed.

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