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During the postgenomics era the information about genomic sequence and gene functions provides a new foundation for evolutionary biology and ecology As the first whole-genome sequenced plant Arabidopsis thaliana and its wild relatives have played a critical role in understanding the evolution of genomics and speciation Both A halleri and A lyrata are closely related to the model species A thaliana A halleri ssp gemmifera occurs in northeastern China Japan and Taiwan; while its sister A halleri ssp halleri is mainly distributed in Europe Geographical barriers such as Tienshan Mountain Range isolate these intraspecific sisters Likewise A lyrata ssp kamchatica and ssp lyrata occur in East Asia and North America respectively Such distribution patterns seem to be consistent with allopartic speciation The comparison between ancestral and extant polymorphism by multilocus can be informative about the population genetics of speciation In this study we collected and analyzed DNA sequences of 98 genes from four wild relatives of A thaliana A halleri ssp gemmifera A halleri ssp halleri A lyrata ssp kamchatica and A lyrata ssp lyrata The ancestral states of these four species were compared to each other in terms of level of genetic variation However the ancestral species at the time of speciation were substantially more polymorphic than the extant geographical populations The observations are not fully compatible with speciation by strict allopatry At some species pairs parapatric speciation seems more reasonable in speciation of Arabidopsis The 98 gene sequences are also used for the congruence test between gene genealogy and species phylogeny Only 28 genes support the species phylogeny but there are 23 genes supports another major genealogy { lyrata} thaliana Based on the phylogenetic position change of A lyrata ssp kamchatica and Ks value for each species pair suggested the recent directional gene flow between A halleri ssp gemmifera and A lyrata ssp kamchatica

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The subject of the full text builds up and divides six major chapters together: Chapter 1 is the foreword, and has studied the current situation and refined and summarized to value and meaning of the selected title background of the thesis, selected title abroad synthetically; Chapter 2, have carried on science to define to the intension of responsibility and governments relevant concepts of responsibility, have explained that constructs the necessity of the responsibility government and realistic meaning; Chapter 3, ility the intension made of the responsibility has been carried on science solves and constructs, summarize the political function and basic principle of administration asking responsibility system, and the logic relation between administration asking and the responsibility government of the responsibility has been analysed scientifically; Chapter 4, has used the administrations of western countries for reference and administration asking responsibility system and high-ranking official asking system of Hong Kong area of our country and the successful experience, expect to the institutional improvement of the administration asking responsibility system of our country can be beneficial; The whole chapter 5, the current administration asking system of our country that has implemented the state and carried on objective analysis in responsibility system, have pointed out a series of questions and not complete places exposed in institutional improvement, and has carried on network analysis from two visual angles of the traditional political culture ecology and system factor aspect to the reason why the question emerges; Chapter 6, the responsibility system has carried on tentative countermeasure analysis and discussion in perfecting the administration of our country, separately from improving the environment of administration asking responsibility and reinventing the culture of administration asking responsibility, devoting more efforts to asking the responsibility in allosome, strengthen the related institutional improvement of administration asking responsibility system, etc.

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