Timber tree of central and southeastern United States having hairy branchlets and a swollen trunk base.

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Leaves evergreen, needlelike, 2-5(-7) per bundle, terminal on very short branchlets, base enclosed by sheath; seed cones maturing in 2nd or 3rd year; seed scales woody, thickened, apex differentiated into apophysis and umbo

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Petals as many as sepals, free, thickened glands on base of adaxial surface.

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Stamen whorls ± equal in shape and length or only slightly unequal in length; anthers subulate to oblong-linear, base tuberculate, expanded, or not appendaged; connective decurrent, forming a short spur.

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Leaves with base cuneate to rounded, or if cordate then apex rounded; leaf blade usually broadest at or above middle.

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Leaves arising from rhizome, simple, alternate, floating, emersed, or submersed, long petiolate but short petiolate on submersed vernal ones; leaf blade undivided, usually with a basal sinus, often peltate.

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Stem fistulose, erect, branched above, base clothed in purplish-brown remnant sheaths.

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Culms flexuose, 3–10 m long, 0.5–1.1 cm in diam., apically clambering; internodes to 60 cm, smooth and glabrous near base, otherwise white powdery and white strigose; wall ca. 1 mm thick. Branches ca.

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No callus could be induced from leaf or petiole explants of varieties F30 and Ⅱ cultured on MS medium if supplemented alone with 6-bezyladenine （6-BA） or thidiazuron, or kinetin. However, the calli were induced on MS medium containing 2.0 mg L-1 6-BA and 0.5 mg L-1 NAA （α-naphthaleneacetic acid）, and the calli derived from petioles of variety Ⅱ could produce redifferentiated shoots, but of variety F30 could not. The later could only produce shoots directly from the base of the petioles.

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Stems stoloniferous, rooting from nodes; leaves 2-4 cm, base broadly cuneate; pedicels 5-6 mm.

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