Plants hirsute with spreading trichomes to 3 mm; fruit (2.5-)3-5 cm × 1.4-2 mm; seeds to 2.5 mm, margined or winged; cotyledons accumbent

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Nutlets 4-5 mm; disc with keel of small anchorlike spines and tuberculate, margin with flat whitish yellow glochids 2-3 mm; glochids 4-6 on each side of disc.

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Fruit dehiscent siliques or rarely silicles, linear or rarely oblong, terete, 4-angled, latiseptate, or angustiseptate, sessile or rarely shortly stipitate; valves with an obscure to prominent midvein, pubescent on outside, rarely also on inside, keeled or not, smooth or torulose; replum rounded; septum complete, membranous, translucent or opaque, veinless; style obsolete or short, rarely half as long as or subequaling fruit, often pubescent; stigma capitate, entire or 2-lobed.

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Fruit dehiscent silicles, oblong, ovoid, ellipsoid, or globose, terete, rarely slightly latiseptate or angustiseptate, sessile; valves veinless or obscurely veined, glabrous, papillate, or with simple trichomes, smooth; replum rounded; septum complete, perforated, or absent; style obsolete or to 3 mm; stigma discoid, entire.

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Ovary superior, 2-loculed; ovules 2 per locule; placentation axile, appearing nearly basal, apotropous or anatropous, bitegmic, crassinucellar; style simple, connate; stigma discoid or capitate, rarely 4-lobed, not papillate.

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Young branches with appressed stellate hairs; leaves 7-14 cm, margin denticulate; petiole more than 1 cm, with glands at apex

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Panicle spikelike, dense, linear to ovate or oblong in outline, lower part sometimes interrupted, 1.5–11 cm; branches short, appressed, pubescent to tomentose. Spikelets 4–9 mm, florets 2(or 3); rachilla hairs 1–1.5 mm; glumes subequal or slightly unequal, lower glume 4–8 mm, upper glume 5–9 mm, apex acuminate, occasionally briefly aristulate; lemmas lanceolate, 4–7 mm, scaberulous to pubescent, awned from upper 1/4–1/3, apex usually 2-denticulate, teeth often mucronate, occasionally subentire; awn 2–7 mm, weakly geniculate with loosely twisted column, or recurved at base, or almost straight; palea keels scaberulous.

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Sterile spikelet ovate, composed of up to 18 stiff, narrowly linear, shortly awned empty lemmas with green ciliate keel.

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Spikelets similar, lanceolate, dorsally compressed; callus bearded with hairs shorter than, as long as, or longer than the spikelet; glumes papery or membranous; lower floret usually represented by a hyaline sterile lemma; upper floret bisexual, lemma hyaline, awned or awnless.

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Glumes (1–)3(–5), transitional with basal, sterile lemmas; lemma 7- to many veined, abaxially commonly puberulent and with distinct tessellation, adaxially slightly scabrous with minute setulae, apex acuminate with an awnlike point; palea conspicuously smaller than lemma or equaling in length but much narrower in upper florets, abaxially minutely setulate, 2-keeled, 2–4-veined between and beside keels, apex obtuse or emarginate; lodicules 3, membranous with thickened lower portion, 5–7-veined, margins ciliate, anterior pair of lodicules usually asymmetrical.

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