Spikelets laterally compressed, florets several, all bisexual or lowest sterile and without a palea; rachilla glabrous, disarticulating above glumes and sterile floret and between fertile florets; glumes lanceolate, shorter than lemmas, subequal or unequal, scarious-membranous, 1–3-veined, obtuse to acuminate or aristulate; lemmas ovate-lanceolate, scarious-membranous, 3-veined, long-ciliate on lateral veins, keeled, apex setaceously bidentate with a short, often recurved awn from the sinus; palea shorter than lemma, hyaline, keels very shortly ciliate.

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The spindle density of collilongus and splenius of head was determined and the expressions of the NT3 and Trk C in dorsal root ganglions were observed immunohistologically.

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Petiole to 8 cm, brown villous and white puberulent; leaf blade subovate, 3-4.5 X 2-3.2 cm, appressed pubescent, abaxially brown villous along veins, margin coarsely crenate-dentate.

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Male flowers (3-)5-7 in cymules, pedicellate; bracts triangular, pilose to ciliate, at base with a pair of elongate-spheroidal to elliptic glands touching axis of thyrse and sometimes decurrent; calyx with 3 sepals, fused at base; petals and disk absent; stamens 3; filament and anther of similar length.

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Sepals and petals 5, persistent after anthesis, sepals free, entire or rarely glandular-ciliate, petals entire or glandular-ciliate, apiculus absent.

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Fruit juicy or dry, naked, scaly, hairy, bristly, or spiny, indehiscent or dehiscent, when juicy then pulp derived from often deliquescent funicles. Seeds usually numerous, often arillate or strophiolate; embryo curved or rarely straight; endosperm present or absent; cotyledons reduced or vestigial, rarely leaflike.

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Spikelets all alike, somewhat laterally compressed, with 1 to several fertile florets; rachilla usually disarticulating below each floret, occasionally strictly 3 florets with lower 2 staminate or barren and of different appearance from upper fertile floret, then disarticulating above glumes and florets falling together; glumes persistent, often equal to spikelet or at least longer than first floret, commonly membranous with broad, shining, hyaline margins; floret callus bearded; lemmas hyaline to leathery, 3- to several-veined ( Coleanthus 1-veined), apex entire or denticulate, awned from back, rarely awnless; awn usually geniculate with twisted column; palea hyaline, subequaling or much shorter than lemma.

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Styles (2 or)3[-5], free. Capsule valves obscurely longitudinally vittate; placentation parietal; seeds not carinate or winged, testa reticulate to ribbed-scalariform.

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Petals obovate or obovate-oblong, 1.1-1.5 cm, ciliolate and brown-glandular on upper half of margin, narrowly clawed at base; petal-scale indistinct, ca. 2 mm, base cuneate, apex truncate and denticulate.

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Spikes few to many flowered; bracts brownish yellow to blackish brown, dull or shining, conchiform, rigid, abaxially minutely papillose toward apex when young, midvein conspicuous, margin scarious, entire, ciliate, fimbriate, or lacerate, apex emarginate, rounded, or acute. Sepals 3; lateral sepals with keel entire, dentate, or ciliate, wings membranous; median sepal enclosing corolla or sometimes ± reduced or obsolete, thin and membranous, 1--3(--5)-veined.

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