Fruit ovoid, slightly dorsally compressed, constricted at the commissure; dorsal and intermediate ribs filiform, lateral ribs obscure, surface glabrous or pilose; vittae 1–2 or 3–6 in each furrow, 2 or 4–8 on commissure.

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Basal leaf blade reniform, rarely ovate-cordate, apex rounded, rarely subobtuse or emarginate; cauline leaf about as large as basal ones; petals obovate, eglandular, apex emarginate

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Racemes axillary or terminal, appendage lamellate; unwinged part of capsule without obvious concentric ribs; seeds ovoid or hirsute-villous, not strophiolate, or pubescent and strophiolate

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Basal leaf blade ovate-cordate; cauline leaf borne near base of stem; staminode lobes oblong.

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Basal leaf blade reniform, rarely ovate-cordate; cauline leaf borne near middle of stem; staminode lobes lanceolate

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Leaves 10-20, red, spirally arranged, imbricate, scaly, longitudinally striate when dry; basal leaves on scape broadly ovate, ca. 1 cm; middle leaves oblong, ca. 2 cm; apical leaves subspatulate, 3-3.5 cm, subtending inflorescence.

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Leaves simple, rarely trifoliolate, opposite, sessile or shortly petiolate, no more than 5 mm; blade ovate to oblong-ovate, 11-14(-16.5)× 5.5-7(-8) cm, coriaceous, glabrous, adaxially lustrous green, abaxially pale green, main vein impressed adaxially and prominent abaxially, lateral veins 5-9(-11) pairs, slightly discernible adaxially and conspicuous abaxially, base subcordate, rarely rounded, margin glandular crenate or -serrate, apex apiculate, cusp 5-10(-15) mm.

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Leaves opposite, usually in many pairs; stipules small; petioles short, connate at base, forming a short sheath; leaf blade elliptic, ovate-elliptic, or elliptic-lanceolate, margin serrate, with 1 gland at serrate tip.

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Stems caespitose, branched or not; leaf blade linear-lanceolate or elliptic-lanceolate, 3-5 mm wide [in Nepal, leaves to 8 mm wide]; racemes terminal or axillary; bracts linear, persistent at flowering; inner sepals ovate-oblong to elliptic-oblong.

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Basal leaves with petioles (0.5-)1.5-4 cm, ciliate with simple trichomes; leaf blade 1- or 2-pinnatesect, oblong or ovate in outline, 1.5-6 × 0.7-3 cm; ultimate segments linear-oblong, oblong, or rarely ovate, 1.5-15 × 0.5-3 mm, apex obtuse or subacute.

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