Leaves alternate, simple; leaf blade reniform to ovate, papery, margin entire.

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Cauline leaves reniform, ovate or triangular, alternate

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Basal leaves petiolate; leaf blade spatulate to ovate, 2-5 cm including petiole, thin, margin sparsely serrate.

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Branches terminating in a large bud, rarely spinescent; leaf blade broadly elliptic or ovate-oblong.

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Ovary usually sessile or slightly stipitate, ovoid, 1-loculed; style terminal, short; stigma capitate.

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Petiole 2-12 mm, white villous; leaf blade ovate to triangular-cordate, 2.1-6 × 1.5-4.2 cm, adaxially sparsely white pubescent, abaxially gray and densely white pubescent, base cordate to subtruncate, margin crenate-serrate, apex acute or rarely obtuse.

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The biodegradation behaviors of six kinds of fatliquors have been researched including sulphated castor sulphated neatsfoot oil, sulphated fish oils, lecithin, chlorinated paraffin and alkyl sulfonyl chloride.

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Fruit ovoid-oblong, slightly compressed laterally, glabrous; ribs 5, filiform; vittae 3–4 in each furrow, 8 on commissure.

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Fruit ovoid-ellipsoid, 1.7–2 × 1–1.5 mm, base cordate, surface granulate with minute papillae; vittae 1 in each furrow, 2 on commissure.

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Fruit ovoid, slightly compressed dorsally; ribs thick-obtuse, very prominent, irregularly denticulate especially along ribs, denticles stiff-membranous or with stiffly scarious-processes and hairs; furrows narrow; vittae 1 in each furrow, 2 on commissure.

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