Flower buds terminal or axillary, ovoid-globose, pubescent with gray or brown appressed trichomes.

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Leaflets narrowly ovate, ovate-lanceolate, or oblong-elliptic, margin sparsely spinose-serrate, both surfaces glabrous, or only crinite at axils of lateral veins.

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Leaves alternate, rarely opposite, petiolate or subsessile; leaf blade flattened, slightly succulent, linear, lanceolate, oblong, ovate, triangular, rhombic, or hastate, margin serrate, rarely entire.

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Leaves paripinnate, alternate, 25-40 cm with petioles; axis and petioles grayish yellow tomentose; leaflets usually 4 pairs, opposite; petiolules 3-4 mm, densely tomentose; blades adaxially deep green, oblong or ovate-oblong, 13-30 × 5-11 cm, papery, abaxially sparsely tomentose along midveins and lateral veins, adaxially sparsely tomentose along midveins, or glabrous on both sides, lateral veins 17-22 pairs, prominent on both sides when dry, base slightly cordate, margin entire, apex caudate-acuminate. Inflorescence racemose or thyrsoid, on stems and old branches, 1.5-4 cm, densely grayish yellow to grayish brown pubescent.

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Basal leaf blade reniform, orbicular, broadly ovate-orbicular, ovate-cordate, or occasionally other shapes.

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Stipular spines erect, 4-17 mm, 2 per node; petiole 5-9 mm, pubescent; leaf blade broadly ovate or ovate-elliptic, 3-5.5(-7)× 2.2-5 cm, papery, abaxially yellowish to brownish pubescent on major veins, adaxially mostly glabrous, distinctly 3-veined from base, base broadly cuneate, cuneate, or subrounded, symmetric to slightly oblique, margin obtusely serrate or serrulate, rarely subentire, apex obtuse, rounded, or slightly retuse. Flowers in axillary cymes.

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Stem leaves opposite, petiolate; leaf blade obovate-spatulate to subrounded, 1-2.5 cm including petiole, margin crenate to subentire.

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Leaves alternate, long petiolate, 2-4 × ternately compound, rarely simple; stipules membranous; leaflets lanceolate, ovate, or broadly ovate to elliptic, margin dentate.

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Fruit ovoid-oblong, slightly flattened laterally, mericarps subrounded in cross section, commissure constricted, glabrous, shiny; ribs 5, filiform, prominent; vittae 1 in each furrow, 2 on commissure.

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Leaves 3, in a terminal whorl, sessile or shortly petiolate, rhombic-orbicular to ovate, with 3 or 5 main veins and anastomosing veinlets.

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