Flowers bisexual or bisexual and male; stamens more than 2 × as many as petals; gynoecium 2- to many loculed; ovules 2 to many per locule; fruit with stalked or rarely sessile pulp vesicles; leaves odd-pinnately 3(or 5)-foliolate, digitately 3-foliolate

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Dioecious monoecious, reproductive structures axillary, solitary. Pollen cones pedicellate, globose, with overlapping bracts at base; microsporophylls 6-14, shield-shaped, each with 4-9 radially arranged pollen sacs. Seed-bearing structures subsessile, with several overlapping bracts basically. Ovule erected, entrusted with disc-like embryophore. After fertilization, embryophore developed into fleshy, cup-like and red aril.

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Male flowers: perianth tube basally solid; stamens 20, connectives completely united to form a stalked cup; anthers inserted on outside at base of cup, surrounding column base; gynostemium rudimentary.

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Inflorescence spikelike, cylindrical, composed of spiny or bristly deciduous burrs arranged along an angular, often sinuous rachis; burrs sessile or with an obconical basal stipe, each composed of 1 or more sessile spikelets surrounded by an involucre of spines and bristles; bristles flexuous or more often spinous,± flattened, grooved on the outer face, united below, the degree of union varying from a small basal disk to a deep cupule, inner spines or bristles often ciliate around spikelets. Spikelets lanceolate, acute; glumes unequal, shorter than spikelet, lower sometimes suppressed; lower floret membranous, staminate or neuter; upper floret firmer, protogynous.

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Female flowers: perianth tube adnate to ovary, mouth contracted, apex 3- or 4-toothed; staminodes absent; stigma sessile, discoid or peltate, long ciliate hairs along margin; ovule orthotropous.

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Male flowers small, 1-3 per bract-axil, subsessile; sepals 3, free or very indistinctly connate at base; petals and disk absent; stamens 3; filaments free; anthers longitudinally dehiscent; pistillode absent.

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Stamens 6. Ovary stalked; style 1; stigmas 3, plumose.

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Lodicules 3. Stamens 6; filaments free. Ovary stalked; style long, hollow; stigmas 2 or 3, plumose.

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Stamens 6; filaments free. Ovary usually stalked, apex thickened and hairy; style solid, usually short; stigmas (1–)3, long, hairy, plumose.

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Staminodes minute, squamose, sessile, inserted at base of carpels opposite petals.

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